4.1. Key Findings
Greater abundance of herbaceous vegetation, forests, and wetlands and low abundance of crops, pasture, and developed open space (lawns, parks, and golf courses) generally explained large fire occurrence (wildfires ≥200 ha), with pronounced regional differences and less distinctive ecoregional differences. More specifically, herbaceous vegetation and herbaceous wetlands and evergreen forest classes were the primary wildland classes that carried fire. These results may help explain the lack of few large fires in 14 states, which contained limited herbaceous vegetation, either as grasslands or wetlands, and an abundance of crops or developed areas, although related factors such as fragmentation from small land parcels likely are influential in preventing fires. Herbaceous vegetation was important for fires in all regions, but in the eastern region, herbaceous vegetation was present in the form of emergent herbaceous wetlands instead of upland grasslands (
Table 1). Although wetlands may be a firebreak when water is present, the herbaceous component of emergent herbaceous wetlands in the eastern region appears to be fire-prone during dry events. Wetlands were less likely to have fires in the central and western regions. Forests overall were more likely to have fires in all regions due to evergreen forests and not mixed or deciduous forests, which were less likely to have fires where these forest types were more influential variables; that is, in the eastern region and southeastern and desert ecoregions. Evergreen forests with low tree densities and an herbaceous layer, which typically result in low severity fires, occur as widespread ponderosa pine (
Pinus ponderosa) in the western region and now relatively rare longleaf and shortleaf pine (
Pinus palustris and echinata) forests and commercial pine plantations in the eastern region [
8]. More densely spaced evergreen species such as spruces, firs, and eastern redcedar, which also have low branches that provide ladder fuels to carry fire vertically to crowns, typically generate higher severity crown fires [
9].
Crops and pasture were the primary land classes where fire did not occur, with developed open space as a less abundant class that also appeared to prevent fire spread. These land classes can be difficult to separate from herbaceous vegetation using remote sensing methods, but differentiation was very clear in where fires did and did not occur (
Table 1). Although these land classes can have fires, fires often are prescribed and small to remove vegetation residue, with limited fuels available to spread fire or escalate fire severity [
2,
28,
29]. Crops, pasture, and developed open space are areas of reduced vegetation, in the gradient between wildlands and developed urban areas with little vegetation. Crops and pasture are harvested or grazed before plants dry and lose enough moisture to ignite easily. Lawns, parks, and golf courses similarly are kept green through watering and low in height through mowing, which limits fuels, and are intersected with impervious surfaces that fragment fuel continuity and prevent fire spread.
Land classes such as crops, pasture, and developed open space with negative influences on fire were as important for where fire did not occur as herbaceous vegetation, forests, and wetlands were greater in abundance where fire did occur, particularly in the eastern and central regions and the southeastern and temperate steppe ecoregions of these regions. In the eastern region, the most influential model variable was a combination of crops, pasture, and developed open space. These land classes covered 32% of the region, 6% of the area where fires occurred, and 38% of the area from samples where fires did not occur. Herbaceous wetlands were among the most important model variables. Herbaceous wetlands were 2% of the region and area from samples where fires did not occur, and 15% of the area where fires occurred. Likewise, in the central region, crops were the most important model variable. Crops were 33% of the region, 2% of the area where fires occurred, and 50% of the area from samples where fires did not occur. Herbaceous vegetation was among the most important model variables, albeit still minor. Herbaceous vegetation was 32% of the area, 72% of the area where fires occurred, and 23% of the area from samples where fires did not occur.
Furthermore, in the central region, forests or evergreen forests, which notably would include eastern redcedar, were not among most influential model variables, unlike grassland herbaceous vegetation that also was more disproportionately represented where fires occurred (2.2 times greater) compared to background land class rates than other wildlands vegetation types (e.g., evergreen forests were greater by a factor of 1.5). By ecoregion, the prairie in particular showed little association of evergreen forests with fire. However, the temperate steppe did contain greater percent area of evergreen forests (5.5%) where fire occurred than background rates (1.5%), with percent area of forests with fires about 2.8-fold greater than background rates. Conversely, herbaceous vegetation was 77% of area where fire occurred compared to 48% of the region, which is a grassland with a great herbaceous vegetation background rate. Nonetheless, the overall wildlands (herbaceous and woody vegetation) and land use (crop and pasture) combinations most influenced fire. These results contrasted with Donovan et al. [
9], who found that woody vegetation burned disproportionately more than other land-use types in the Great Plains. However, Donovan et al. [
9], expected to find that large fires occurred in grasslands, which are more fire-prone than woodlands.
The shrub land class was minimally influential on fire, but perhaps that will change due to recent fires in the shrub chaparral of California. Fire-dependent shrub ecosystems also occur in shrubs of the Lake Wales Ridge in Florida and pine barrens of the mid-Atlantic eastern US, and albeit small remnants, these types of ecosystems are examples to explain greater abundance of shrubs where fires occurred in the eastern region. Additionally, increased fire activity occurs because of invasion by primarily non-native cheatgrass in shrublands of the Great Basin in the desert ecoregion and a complex of non-native grass invasions in shrublands of the subtropical ecoregion; non-native grasses provide horizontal fuel continuity in fuel-limited shrub systems [
13]. However, fire then occurs in predominantly herbaceous vegetation rather than shrubs.
Fire relationships were not consistent in space, although the central region shared overlap with both the eastern and western regions. Models for the eastern region overpredicted fires in the western region and likewise, models for the western region underpredicted fires in the eastern region. That is, vegetation that would produce a fire in the eastern region was not sufficient for fire in the western region and vegetation that would not produce a fire in the western region was sufficient for fire in the eastern region. These differences in part may be due to more land classes associated with fires in the eastern region, including wetlands and deciduous forest, than in the western region. These results also seem to reflect that the western region typically has a drier climate, which can result in fuel limitations in vegetation but also greater probability of vegetation being cured and able to ignite for a longer time interval and extreme weather conditions suitable for fire [
3,
30]. Nonetheless, despite greater current rates of fires in the west, historically, the longleaf pine “grasslands” of the Coastal Plain in the southeastern U.S. had fire regimes of 2 to 5 years, making it one of the most frequently burned regions in the past, despite a humid, wet climate [
8,
9]. To a lesser magnitude, predictions among nearby ecoregions had the same problem, in that eastern ecoregional models overpredicted fires for the southeastern ecoregion and the southeastern models underpredicted fire for the eastern ecoregion and the temperate steppe also underpredicted fire for the prairie region.
Fire relationships were not consistent in time. The 1999 to 2002 models were less successful at forecasting 2014 to 2017 fires than 2014 to 2017 models predicted 1999 to 2002 fires. Number of fires greatly increased in the central region from 310 fires during 1999 to 2002 to 1569 fires during 2014 to 2017, while number of fires in the other regions stayed relatively stable. Compared to 1999 to 2002 models, importance of herbaceous and crop land covers increased for the 2014 to 2017 models, which still remained successful at hindcasting 1999 to 2002 fires.
The three classifiers performed near equally well at predicting based on models of the same extent or time interval, or similar ecoregions, despite a variety of potential errors arising from for example, the fire perimeters, land class misclassification, and areas that could have burned yet did not. However, in general, land classes were well-differentiated between where fire occurred and samples of where fire was not known to occur (
Table 1). Predictions across time or space in different regions showed that random forests did not perform as well as the other two classifiers. Mean value of true positive rate for extreme gradient boosting was 0.790 compared 0.761 for random forests when predicting across time intervals. Mean value of true positive rate for extreme gradient boosting was 0.708 compared 0.655 for random forests when predicting across regions, albeit if the random forests prediction for the central region based on the eastern region is excluded, the mean value increases to 0.691. These two examples demonstrate that extreme gradient boosting remained more generalized with less overfitting to specific datasets than random forests, although the sample size is too small to state that extreme gradient boosting overall is preferable to random forests when models are in development or need flexibility. Moreover, random forests performed equally well in predicting fires in adjacent ecoregions, which had fewer differences than across regions.
Analysis with multiple classifiers allows identification of better performing classifiers and selection of desirable features. Random forests is a well-established classifier with reliably accurate performance, whereas C5.0 decision trees generate explicit rulesets, which can be help explain models when there are few variables, but also produce many similar and competing models. Fernandez-Delgado et al. [
18] comprehensively assessed 179 classifiers using 121 data sets and the best classifier was random forests, with C5.0 decision trees among the best classifiers. Extreme gradient boosting is newer classifier, intended to reduce overfitting that other boosting classifiers are prone to producing [
19].
Further research may explore some unexpected results from these relationships. For example, future studies can examine the interaction of fire with mixed and deciduous forests, which overall were not associated with fires. These forest types were less abundant where fires occurred when these forests were among the most influential model variables; for example, mixed forests in the eastern region and deciduous forests in the southeastern and desert ecoregions. It may be that current species in mixed and deciduous forests are not fire-tolerant, and thus suppress rather than support fires, reducing fire frequency but increasing fire risk when fire does occur [
31,
32]. Furthermore, presence of water in mixed and deciduous forests will act as a firebreak. Deciduous species are rare in the western region and often occur along riparian networks, and likewise, deciduous and mixed forests in the southeastern U.S. often form along wetlands. Similarly, the importance of herbaceous wetlands, as a substitute for herbaceous vegetation, for fires in the eastern region also may be assessed to explore the interaction between the fire-spreading herbaceous component and the fire-breaking wetlands component.
Although this study provides summaries that clearly show fires are more prevalent in some land classes than others, varying be region, along with accurate predictions of models at least within modeling extents, limitations include errors in data and also that land cover is just one of many factors that influence fire. Fires vary in severity and include unburned areas within perimeters, fires may not be detected, land classes may be misidentified, and most areas that are flammable did not burn during the time interval of this study [
5]. Fires also are influenced by time since last fire, vegetation properties in terms of acting as fuels, weather and climate, number of ignitions, and topography. Additionally, spatial arrangement of land classes also is an important component to whether fire will be able to spread across flammable wildlands vegetation or be limited by interspersed firebreaks that break fuel continuity. Furthermore, fire protection services in conjunction with wildlands fragmentation and lack of extreme weather may suppress fires while they are small. For example, some of the 14 states with few large fires, particularly New York, indeed have many ignitions, resulting in numerous fires <200 ha [
23,
33]. Land management through prescribed burns also will reduce fuels and reduce the risk of fire.
4.2. Implications
Fire is a fundamental ecological process that maintains ecosystems and associated species by filtering species that can survive fire or reproduce in response to fire. Fire favors herbaceous vegetation, which rapidly captures aboveground growing space, over woody vegetation, and thereby supports a diverse and unique assemblage of species, including insects that provide many critical ecosystem services [
34,
35,
36,
37]. Hominins have applied fire as a labor-saving land management tool for hundreds of thousands of years [
38]. However, as human densities have increased, fire has become a risk to human structures and lives. Fire had been suppressed for about a century in the United States, resulting in great departure from historical fire regimes. Nonetheless, these models demonstrated that fires still follow historical ecology that herbaceous vegetation and pine forests are fire-prone, with traits that promote fire [
8].
Although fire suppression appears to be a logical response to fire, fire ecology produces a paradox that fires become more catastrophic because of, not despite, fire suppression [
14,
39,
40]. Fuels accumulate, increasing in quantity and continuity, when fuels do not burn due to suppression of unintended fires, exclusion of prescribed fires for land management, along with passive suppression from land classes that act as fire breaks (described in this study). Changing characteristics of fires often culminates in disastrous fires that cannot be suppressed [
14]. Fire suppression produces an escalating cycle of need for fire suppression and increased fire suppression costs [
14]. This aspect of fire ecology has been understood for over a century. For example, prescribed burns were established practice during the late 1800s in California forests to prevent high severity crown fires [
41]. Show and Kotok (1924, [
41]) wrote,
The idea that fires could be excluded entirely from millions of acres was generally regarded as preposterous and the most gloomy pictures were drawn of any such attempt. It was claimed that the uncontrollable crown fire was to be expected as the inevitable consequence of allowing ground cover and litter to accumulate. Thus, in the early years of protection of the national forests, the forests were still open as a result of the repeated fires of the past…. As fire protection became an accomplished fact and the young growth began to fill up the open forest, the amount of inflammable material in the forests increased greatly.
Compounding altered fire regimes, housing developments and other infrastructure have expanded into wildlands area, increasing the area of wildland-urban interface (WUI) and intermix and concurrently, exposure of assets to fire throughout the world and resulting in structural losses to fire events [
14]. Fire severity in wildlands may be relatively independent of fire in the home ignition zone; that is, homeowner management of the area immediately surrounding the determines home losses from wildland fire [
14]. Thus, in addition to untreated fuel accumulation in wildlands and increased exposure and ignition rates due to development in wildlands, many residents in wildlands underinvest in mitigation actions, relying on neighboring public or private land owners for fuel treatment, government emergency services to suppress fire, and insurance or federal disaster assistance to cover any property losses [
42,
43]. Housing materials, such as wood shakes, can be very flammable and burning homes generate hot embers, which increase the spread of a fire. Lofted embers may cause more residential losses than direct flames or radiant heat [
14].
Strategies for risk mitigation to coexist with fire include identification of communities at risk, disclosure of fire risk, reduction of vegetation (i.e., fuel for fire), outreach to homeowners to reduce both structure flammability and vegetation surrounding structures, and planning and zoning to reduce sprawl into wildlands [
33]. For an example of outreach and risk disclosure, counties can provide advisory letters informing citizens about risks and homeowner responsibilities, similar to the following, “The physical characteristics of your property can be positive and negative. Trees are a wonderful environmental amenity, but can also involve your home in a forest fire. Building at the top of a forested draw should be considered as dangerous as building in a flash flood area. ‘Defensible perimeters’ are very helpful in protecting buildings from forest fire and inversely can protect the forest from igniting if your house catches on fire.” (country commissioners in Stevens County, MT, USA, as cited by [
44]). Reduction of intermixing between wildlands and housing, by concentration of housing at greater densities, will reduce fire exposure. These models show that creative planning and zoning may allow placement of firebreak buffer zones of crops, grazed pastures, and open spaces that are managed by mowing and watering between wildlands and developed land classes with housing.