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Quaternary, Volume 7, Issue 4 (December 2024) – 12 articles

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25 pages, 11533 KiB  
Article
Hydroclimatic Changes Revealed by Multiple Proxies Since the Last Glacial Maximum from the Core Monsoon Zone of India
by Mohammad Firoze Quamar, Jyotsna Dubey, Pooja Tiwari, Prasanta Kumar Das, Biswajeet Thakur, Mohammad Javed, Nagendra Prasad, M. E. T. Maneesha and Satish J. Sangode
Quaternary 2024, 7(4), 52; https://doi.org/10.3390/quat7040052 - 21 Nov 2024
Viewed by 873
Abstract
We present multiproxy records from a 2.25-m-long lake sediment profile from central India, which suggested that between ~22,200 and 18,658 cal yr BP, the Indian Summer Monsoon (ISM) was weak, supporting open vegetation in a cool and dry climate, which is globally correlated [...] Read more.
We present multiproxy records from a 2.25-m-long lake sediment profile from central India, which suggested that between ~22,200 and 18,658 cal yr BP, the Indian Summer Monsoon (ISM) was weak, supporting open vegetation in a cool and dry climate, which is globally correlated with the Last Glacial Maximum (LGM). The grain size data of this phase suggest low-energy conditions, indicating a weak ISM. Environmental magnetic concentration-dependent parameters also confirm this weakened ISM. Between ~18,658 and 7340 cal yr BP, the ISM underwent a notable increase, and open mixed tropical deciduous forests replaced the existing vegetation under a warm and moderately humid climate. Environmental magnetic parameters and the grain size data signal a shift toward higher energy levels, in harmony with the warm and moderately humid climate during this time span. Between ~7340 and 1960 cal yr BP, the ISM intensity further increased, which supported open mixed tropical deciduous forests with a rise in prominent tree species under a warm and a relatively more humid climate, correlated with the global Holocene Climatic Optimum (HCO). The trends in environmental magnetic parameters and grain size data mirror this phase of climatic amelioration. From ~1961 cal yr BP to the present, the ISM has intensified, giving rise to dense mixed tropical deciduous forests under a warm and relatively more humid climate. Environmental magnetic parameters and the grain size data are in tandem with the palynogical findings from this phase of the ISM variability. Full article
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8 pages, 290 KiB  
Comment
The Late Villafranchian Absence of Pigs in Europe. Comment on Iannucci, A. The Occurrence of Suids in the Post-Olduvai to Pre-Jaramillo Pleistocene of Europe and Implications for Late Villafranchian Biochronology and Faunal Dynamics. Quaternary 2024, 7, 11
by Bienvenido Martínez-Navarro, Joan Madurell-Malapeira, Sergio Ros-Montoya, M. Patrocinio Espigares, Guillermo Rodríguez-Gómez, Lorenzo Rook and Paul Palmqvist
Quaternary 2024, 7(4), 51; https://doi.org/10.3390/quat7040051 - 21 Nov 2024
Viewed by 255
Abstract
On 2015, after the direct study of the most important Late Villafranchian fossil collections of Europe and Western Asia, including Orce (Spain), Pirro Nord and Upper Valdarno (Italy), Appollonia (Greece), Dmanisi (Georgia) and ‘Ubeidiya (Israel), among others, our team proposed the hypothesis that [...] Read more.
On 2015, after the direct study of the most important Late Villafranchian fossil collections of Europe and Western Asia, including Orce (Spain), Pirro Nord and Upper Valdarno (Italy), Appollonia (Greece), Dmanisi (Georgia) and ‘Ubeidiya (Israel), among others, our team proposed the hypothesis that suids disappeared from Europe during the time span between 1.8 and 1.2 Ma. The implications of our conclusions were significant, the arrival of Early Homo into Western Europe, dated to 1.4 Ma at the site of Barranco León in Orce (Spain), preceded the return of pigs into the continent at 1.2 Ma. This hypothesis has been recently challenged because of the finding of an incomplete metatarsal ascribed to Sus sp., with no clear stratigraphic origin, found in the XIX Century Croizet collection of Peyrolles (France), which is housed in the Natural History Museum, London, together with other weak arguments based on the absence of reliable dating for many Early Pleistocene European sites, and other hypothetical records of pigs, with no real fossil support. We answer all these questions and defend that our 2015 hypothesis is correct. Full article
22 pages, 6306 KiB  
Article
The Evolution of Long-Range Hunting with Stone-Tipped Weapons During the Afrotropic Middle Stone Age: A Testable Framework Based on Tip Cross-Sectional Area
by Yonatan Sahle and Marlize Lombard
Quaternary 2024, 7(4), 50; https://doi.org/10.3390/quat7040050 - 18 Nov 2024
Viewed by 486
Abstract
In the Afrotropic biogeographic realm, with its diverse and high-density mammal population, early humans may have been hunting with stone-tipped weapons since ~500,000 years ago. Being able to hunt effectively from a distance has several important adaptive advantages. Yet, until now, African long-range [...] Read more.
In the Afrotropic biogeographic realm, with its diverse and high-density mammal population, early humans may have been hunting with stone-tipped weapons since ~500,000 years ago. Being able to hunt effectively from a distance has several important adaptive advantages. Yet, until now, African long-range javelin hunting remained unexplored as intermediate between short/medium-range, hand-delivered and long-range, mechanically projected weapons. Insights gained from a new Afrotropic comparative dataset with 950 weapon tips of known use—including several javelin types—provide a contextually appropriate middle-range tool for assessing the probable effective hunting ranges of Middle Stone Age points. We use a novel application of the ballistically relevant tip cross-sectional area (TCSA) statistic to define contact, short-, medium-, long- and maximum-range hunting and discuss the adaptive advantages for each. The approach is applied to suggest developments and variations in the best-fit hunting ranges of 5597 stone points from 62 Middle Stone Age Afrotropic assemblages. By aligning our results with the Marine Isotope Stage (MIS) record we hypothesize that effective long-range (~20–30 m) hunting with stone-tipped weapons was probably not practiced by ≥MIS 8, and that experimentation with long-range javelins—similar to those used by contemporary Ethiopian hunters—over these distances may have started during MIS 6, becoming part of the everyday Afrotropic hunting arsenal by the end of MIS 5. Full article
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19 pages, 3429 KiB  
Article
Were Neanderthals the First Collectors? First Evidence Recovered in Level 4 of the Prado Vargas Cave, Cornejo, Burgos and Spain
by Marta Navazo Ruiz, Alfonso Benito-Calvo, María Carmen Lozano-Francisco, Rodrigo Alonso Alcalde, Pedro Alonso García, Héctor de la Fuente Juez, Marta Santamaría Diez and Paula Cristóbal Cubillo
Quaternary 2024, 7(4), 49; https://doi.org/10.3390/quat7040049 - 12 Nov 2024
Viewed by 6835
Abstract
Collecting is a form of leisure, and even a passion, consisting of collecting, preserving and displaying objects. When we look for its origin in the literature, we are taken back to “the appearance of writing and the fixing of knowledge”, specifically with the [...] Read more.
Collecting is a form of leisure, and even a passion, consisting of collecting, preserving and displaying objects. When we look for its origin in the literature, we are taken back to “the appearance of writing and the fixing of knowledge”, specifically with the Assyrian King Ashurbanipal (7th century BC, Mesopotamia), and his fondness for collecting books, which in his case were in the form of clay tablets. This is not, however, a true reflection, for we have evidence of much earlier collectors. The curiosity and interest in keeping stones or fossils of different colors and shapes, as manuports, is as old as we are. For decades we have had evidence of objects of no utilitarian value in Neanderthal homes. Several European sites have shown that these Neanderthal groups treasured objects that attracted their attention. On some occasions, these objects may have been modified to make a personal ornament and may even have been integrated into subsistence activities such as grinders or hammers. Normally, one or two such specimens are found but, to date, no Neanderthal cave or camp has yielded as many as the N4 level of Prado Vargas Cave. In the N4 Mousterian level of Prado Vargas, 15 specimens of Upper Cretaceous marine fossils belonging to the Gryphaeidae, Pectinidae, Cardiidae, Pholadomyidae, Pleurotomariidae, Tylostomatidae and Diplopodiidae families were found in the context of clay and autochthonous cave sediments. During MIS 3, a group of Neanderthals transported at least fifteen marine fossils, which were collected from various Cretaceous units located in the surrounding area, to the Prado Vargas cave. The fossils, with one exception, show no evidence of having been used as tools; thus, their presence in the cave could be attributed to collecting activities. These activities could have been motivated by numerous tangible and intangible causes, which suggest that collecting activities and the associated abstract thinking were present in Neanderthals before the arrival of modern humans. Full article
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23 pages, 2662 KiB  
Review
Old and New Approaches in Rock Art: Using Animal Motifs to Identify Palaeohabitats
by Mirte Korpershoek, Sally C. Reynolds, Marcin Budka and Philip Riris
Quaternary 2024, 7(4), 48; https://doi.org/10.3390/quat7040048 - 7 Nov 2024
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Abstract
Humans are well known to have made paintings and engravings on rock surfaces, both geometric motifs with an unclear representation, and representative motifs that refer to their activities and aspects of their environment. This kind of art is widespread across time and space [...] Read more.
Humans are well known to have made paintings and engravings on rock surfaces, both geometric motifs with an unclear representation, and representative motifs that refer to their activities and aspects of their environment. This kind of art is widespread across time and space and has throughout history been subjected to various kinds of approaches. Typically, rock art research focuses on its role in the development of the hominin brain and the capability of abstract thinking, as well as on interpreting representative and non-representative motifs. Ethnography and cognitive research have often stressed that rock art is the result of ritual practises and the expression of a shamanic belief system. However, representative motifs may also shed light on a region’s ecological and human prehistory. Here, we give an overview of the general development of rock art study: we highlight the development of artistic behaviour in humans by discussing aesthetic preferences, and the creation of simple geometric motifs and eventually representative motifs, before describing the theories that developed from the earliest study of rock art. These have largely focused on classification and interpretation of the motifs, and often centred on Palaeolithic material from Europe. We then move on to discuss how ethnography among rock art creating communities often suggests important relationships between specific animals in both the realms of spiritual belief systems and within the local environment. Lastly, we highlight how rock art reflects the local penecontemporaneous environment when it comes to depictions of animals, plants, technologies, humans and their activities. We argue that animal depictions are a useful subject to study on a large scale, as it is the most widespread representative motif, and the most appropriate subject to study when the goal is to draw conclusions on environmental changes. Rock art can fill gaps in the local archaeological record and generate new questions of it, but also offer new insights into the history of local human–animal interaction: animal species depicted and/or referred to in rock art are likely to have been a selection of spiritually important animals and a comparison to known information on human interactions with local species may reveal patterns among which animals are selected for local rock art depictions and which are not. Interregional comparison can in turn shed light on whether humans in general tend to ascribe meaning to the same types of animals. We end the review with suggestions for future study, with a special role for computational methods, which are suitable for the analysis of large databases of visual imagery. Full article
(This article belongs to the Special Issue Climate Change versus Cultural Heritage: Past, Present and Future)
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32 pages, 14028 KiB  
Article
A Paleoenvironmental Reconstruction of Lake Vrana on the Island of Cres (Croatia) Based on the Geochemistry and Mineralogy of the Late Pleistocene and Holocene Sediments
by Nikolina Ilijanić, Slobodan Miko, Ozren Hasan, Dea Brunović, Martina Šparica Miko and Saša Mesić
Quaternary 2024, 7(4), 47; https://doi.org/10.3390/quat7040047 - 28 Oct 2024
Viewed by 517
Abstract
A 7.4 m long sediment core has been retrieved from the central part of Lake Vrana on the island of Cres to reconstruct the paleoenvironmental conditions. Lake Vrana is the deepest freshwater lake in Croatia, located in the karst region of the eastern [...] Read more.
A 7.4 m long sediment core has been retrieved from the central part of Lake Vrana on the island of Cres to reconstruct the paleoenvironmental conditions. Lake Vrana is the deepest freshwater lake in Croatia, located in the karst region of the eastern Adriatic coast. A dated sediment sequence in Lake Vrana of 4.4 m has spanned the past 16.4 kyr, and it featured a dynamic sediment deposition until the beginning of the Holocene, including strong sediment input and supply to the lake by runoff sediments of dolomitic origin from the catchment in the period 16.4–14.4 cal kyr BP. High organic carbon content, which originates from mixed terrestrial and aquatic origins in the periods 14.4–13.3 cal kyr BP and 12.7–11.7 cal kyr BP, indicates fluctuating lake levels in shallow water environments during the Late Glacial to Holocene transition. The Holocene sequence indicates the development of more stable conditions and continuous sediment deposition, characterized by an increasing trend of siliciclastic sediments delivered into the lake during the early Holocene (11.7–10 cal kyr BP) and dominantly from 8 to 4.4 cal kyr BP, indicating enhanced input and erosion, which coincides with the humid and pluvial period recorded in the central Mediterranean region. It is followed by sediments with high organic carbon content between 4.4 and 1.6 cal kyr BP, which points to higher lake productivity. Calcite sedimentation prevailed between 1.6 to 0.4 cal kyr BP, indicating stable deeper-lake conditions. Predominantly, siliciclastic sediments from 0.4 to 0.1 cal kyr BP pointed to erosion during the Little Ice Age (LIA), with enhanced precipitation and sediment discharge from the catchment. The re-establishment of calcite sedimentation has been observed over the last 100 years. Full article
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19 pages, 4996 KiB  
Article
Characterization of Heavy Minerals and Their Possible Sources in Quaternary Alluvial and Beach Sediments by an Integration of Microanalytical Data and Spectroscopy (FTIR, Raman and UV-Vis)
by Adel A. Surour and Amira M. El-Tohamy
Quaternary 2024, 7(4), 46; https://doi.org/10.3390/quat7040046 - 22 Oct 2024
Viewed by 586
Abstract
Quaternary stream sediments and beach black sand in north-western Saudi Arabia (namely Wadi Thalbah, Wadi Haramil and Wadi Al Miyah) are characterized by the enrichment of heavy minerals. Concentrates of the heavy minerals in two size fractions (63–125 μm and 125–250 μm) are [...] Read more.
Quaternary stream sediments and beach black sand in north-western Saudi Arabia (namely Wadi Thalbah, Wadi Haramil and Wadi Al Miyah) are characterized by the enrichment of heavy minerals. Concentrates of the heavy minerals in two size fractions (63–125 μm and 125–250 μm) are considered as potential sources of “strategic” accessory minerals. A combination of mineralogical, geochemical and spectroscopic data of opaque and non-opaque minerals is utilized as clues for provenance. ThO2 (up to 17.46 wt%) is correlated with UO2 (up to 7.18 wt%), indicating a possible uranothorite solid solution in zircon. Hafnoan zircon (3.6–5.75 wt% HfO2) is a provenance indicator that indicates a granitic source, mostly highly fractionated granite. In addition, monazite characterizes the same felsic provenance with rare-earth element oxides (La, Ce, Nd and Sm amounting) up to 67.88 wt%. These contents of radionuclides and rare-earth elements assigned the investigated zircon and monazite as “strategic” minerals. In the bulk black sand, V2O5 (up to 0.36 wt%) and ZrO2 (0.57 wt%) are correlated with percentages of magnetite and zircon. Skeletal or star-shaped Ti-magnetite is derived from the basaltic flows. Mn-bearing ilmenite, with up to 5.5 wt% MnO, is derived from the metasediments. The Fourier-transform infrared transmittance (FTIR) spectra indicate lattice vibrational modes of non-opaque silicate heavy minerals, e.g., amphiboles. In addition, the FTIR spectra show O-H vibrational stretching that is related to magnetite and Fe-oxyhydroxides, particularly in the magnetic fraction. Raman data indicate a Verwey transition in the spectrum of magnetite, which is partially replaced by possible ferrite/wüstite during the measurements. The Raman shifts at 223 cm−1 and 460 cm−1 indicate O-Ti-O symmetric stretching vibration and asymmetric stretching vibration of Fe-O bonding in the FeO6 octahedra, respectively. The ultraviolet-visible-near infrared (UV-Vis-NIR) spectra confirm the dominance of ferric iron (Fe3+) as well as some Si4+ transitions of magnetite (226 and 280 nm) in the opaque-rich fractions. Non-opaque heavy silicates such as hornblende and ferrohornblende are responsible for the 192 nm intensity band. Full article
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17 pages, 6063 KiB  
Article
Holocene Climatic Changes in the Negev Desert (Israel): Pollen and Stable Isotopes (13C/12C and 15N/14N) Records from Atzmaut and Ramon I Zoogenic Deposits
by Anna Babenko
Quaternary 2024, 7(4), 45; https://doi.org/10.3390/quat7040045 - 17 Oct 2024
Viewed by 540
Abstract
Two zoogenic deposits from the central part of the Negev Desert (Israel) were investigated by stable isotopes (carbon 13C/12C and nitrogen 15N/14N) and pollen analyses. The merger of these data and results of radiocarbon dating of Atzmaut [...] Read more.
Two zoogenic deposits from the central part of the Negev Desert (Israel) were investigated by stable isotopes (carbon 13C/12C and nitrogen 15N/14N) and pollen analyses. The merger of these data and results of radiocarbon dating of Atzmaut and Ramon I deposits enabled us to reconstruct climate and vegetation changes in the Negev Desert over the past 8500 years. Decrease of the δ13C value in plant remains is a sensitive indicator of paleoclimatic conditions in the region. The decline of the δ13C value over the past 8500 years almost fully coincides with an increase of the total pollen concentration, the proportion of Poaceae pollen in the profile of zoogenic deposits and wetter periods. Thus, four humid periods are identified, from the middle of the 4th millennium to the end of 3rd millennium BC, the 1st half of the 2nd millennium BC, from the 2nd half of the 1st millennium BC to the 1st half of the 1st millennium AD and the middle of the 2nd millennium AD. The δ15N value of plant remains is a less sensitive indicator of climate dynamics and represents the most significant change of precipitation in the region by the end of the 3rd millennium BC. Full article
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21 pages, 8114 KiB  
Article
Palaeoecological Conditions in the South-Eastern and Western Baltic Sea during the Last Millennium
by Ekaterina Ponomarenko, Tatiana Pugacheva and Liubov Kuleshova
Quaternary 2024, 7(4), 44; https://doi.org/10.3390/quat7040044 - 14 Oct 2024
Viewed by 715
Abstract
We present the reconstruction of palaeoenvironmental conditions in the Gdansk, Bornholm, and Arkona Basins of the Baltic Sea over the last millennium. A multiproxy study (including geochemical, XRF, grain size, AMS, and micropalaeontological analyses) of five short sediment cores was performed. The relative [...] Read more.
We present the reconstruction of palaeoenvironmental conditions in the Gdansk, Bornholm, and Arkona Basins of the Baltic Sea over the last millennium. A multiproxy study (including geochemical, XRF, grain size, AMS, and micropalaeontological analyses) of five short sediment cores was performed. The relative age of the sediments was determined based on the Pb distribution along the sediment sequences, as radiocarbon dating has resulted in an excessively old age. The retrieved cores cover two comparable warm periods, the Medieval Climate Anomaly and the Modern Warm Period, for which the increase in surface water productivity was reconstructed. Notably, the production of diatoms was higher during the colder periods (the Dark Ages and Little Ice Age), but this was also the case within the Modern Warm Period. In the Gdansk Basin, the initial salinity increase during the Littorina transgression started after 7.7 cal. a BP. The increased inflow activity was reconstructed during the Medieval Climate Anomaly, even in the Gdansk Basin, despite, in general, very low foraminiferal amounts and diversity. The strongly positive North Atlantic Oscillation Index during this period led to the prevalence of westerly winds over the Baltic region and stronger saltwater intrusions. In the recent sediments, the reconstructed inflow frequency demonstrates a variability against the reduction trend, and a general decline compared to the Medieval Climate Anomaly is seen. Full article
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22 pages, 2208 KiB  
Article
The Final Pliocene and Early Pleistocene Faunal Dispersals from East to Europe and Correlation of the Villafranchian Biochronology between Eastern and Western Europe
by Nikolai Spassov
Quaternary 2024, 7(4), 43; https://doi.org/10.3390/quat7040043 - 11 Oct 2024
Viewed by 911
Abstract
The Villafranchian stage in the mammal fauna evolution in Eurasia (ca. 3.6/3.4 Ma—ca. 1.2 Ma) is associated with the beginning of the formation of the modern appearance of the mammal megafauna of today’s Palaearctic. The cooling and the aridification starting with the beginning [...] Read more.
The Villafranchian stage in the mammal fauna evolution in Eurasia (ca. 3.6/3.4 Ma—ca. 1.2 Ma) is associated with the beginning of the formation of the modern appearance of the mammal megafauna of today’s Palaearctic. The cooling and the aridification starting with the beginning of the Early Pleistocene gradually eliminated the quasi-tropical appearance of the Late Neogene landscapes and fauna of Europe. The time from the Mid-Piacenzian (ca. 3.3–3.0 Ma) to the end of the Early Pleistocene was a time of particularly intense dispersal of species, of faunal exchange between Eurasia and Africa, and of the entry of new mammals into Europe from the East. That is why the correlation of the biochronology of the Villafranchian fauna between Eastern and Western Europe is of particular interest. Accumulated data make possible a more precise correlation of these faunas today. A correlation of selected Eastern European localities with established faunal units and MNQ zones is made in the present work. Usually, the dispersal from Asia or from E. Europe to W. Europe is instantaneous from a geological point of view, but in a number of cases, reaching W. Europe happens later, or some species known to be from Eastern Europe do not reach Western Europe. The main driving forces of the faunal dispersals, which are the key bioevents in the faunal formation, are climate changes, which in turn, affect the environment. We can summarize the following more significant Villafranchian bioevents in Europe: the End Pliocene (Early Villafranchian: MNQ16) turnover related to the first appearance of a number of taxa, for example, felids, canids, proboscideans, and ungulates; the Quaternary beginning turnover. Correlated with this are the beginning of the Middle Villafranchian, which should be placed at about 2.6 Ma; the Coste San Giacomo faunal unit turnover (Senèze and Slivnitsa localities should be included here, and the FU itself, at the very beginning of the late Villafranchian (=MNQ18a)); the Pachycrocuta event at the very beginning of the Olivola FU; and the events related to the Late Villafranchian/Epivillafranchian bounfary. Full article
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11 pages, 4970 KiB  
Article
Plio-Pleistocene Small Mammal-Based Biochronology of Eastern Anatolia and Transcaucasus
by Alexey S. Tesakov, Pavel Frolov, Alexandra Simakova, Albina Yakimova, Vadim Titov, Pranav Ranjan, Hasan Çelik and Vladimir Trifonov
Quaternary 2024, 7(4), 42; https://doi.org/10.3390/quat7040042 - 29 Sep 2024
Viewed by 549
Abstract
The known Plio-Pleistocene mammalian record, mainly represented by small mammals, and its biotic and geological context in the vast region of Eastern Turkey and Transcaucasus provides a sound base for regional biochronology. Recently obtained faunal associations and the main evolutionary lineages found in [...] Read more.
The known Plio-Pleistocene mammalian record, mainly represented by small mammals, and its biotic and geological context in the vast region of Eastern Turkey and Transcaucasus provides a sound base for regional biochronology. Recently obtained faunal associations and the main evolutionary lineages found in the region support direct correlations to the European (ELMA/MN/MQ) and the Eastern European (faunal complexes/MQR-MNR) biochronological systems. Important data on palynology, aquatic and terrestrial mollusks, and magnetostratigraphy integrate the reviewed material into a robust local biochronology. The range of standard biochrons of Early Pliocene through late Early Pleistocene and the regional Anatolian zones M-P are reliably detected. The Early Pleistocene time range (zone P) is refined based on rhizodont lagurines Borsodia and Euro-Asian larger voles Mimomys ex gr. pliocaenicus. The successive zone R for Early Pleistocene faunas with early rootless Microtini is proposed. Full article
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33 pages, 49504 KiB  
Article
The Late Early–Middle Pleistocene Mammal Fauna from the Megalopolis Basin (Peloponnese, Greece) and Its Importance for Biostratigraphy and Paleoenvironment
by George E. Konidaris, Athanassios Athanassiou, Vangelis Tourloukis, Krystalia Chitoglou, Thijs van Kolfschoten, Domenico Giusti, Nicholas Thompson, Georgia Tsartsidou, Effrosyni Roditi, Eleni Panagopoulou, Panagiotis Karkanas and Katerina Harvati
Quaternary 2024, 7(4), 41; https://doi.org/10.3390/quat7040041 - 24 Sep 2024
Viewed by 2090
Abstract
Recent investigations in the upper Lower–Middle Pleistocene deposits of the Megalopolis Basin (Greece) led to the discovery of several sites/findspots with abundant faunal material. Here, we provide an updated overview including new results on the micro- and macro-mammal fauna. Important new discoveries comprise [...] Read more.
Recent investigations in the upper Lower–Middle Pleistocene deposits of the Megalopolis Basin (Greece) led to the discovery of several sites/findspots with abundant faunal material. Here, we provide an updated overview including new results on the micro- and macro-mammal fauna. Important new discoveries comprise partial hippopotamus skeletons from Marathousa 1 and the new Lower Pleistocene site Choremi 6, as well as a second partial elephant skeleton from Marathousa 1, including a complete tusk and the rarely found stylohyoideum. Based on the first results from the newly collected micromammals, we discuss age constraints of the sites, and we provide biostratigraphic/biochronologic remarks on key mammal taxa for the Middle Pleistocene of Greece and southeastern Europe. The presence of mammals highly dependent on freshwater for their survival, together with temperate-adapted ones in several stratigraphic layers of the basin, including those correlated with glacial stages, when conditions were colder and/or drier, indicate the capacity of the basin to retain perennial freshwater bodies under milder climatic conditions, even during the harsher glacial periods of the European Middle Pleistocene, and further support its refugial status. Yet, the smaller dimensions of the Megalopolis hippopotamuses may represent a response to the changing environmental conditions of the epoch, not optimal for hippopotamuses. Overall, the Megalopolis Basin comprises a unique fossil record for southeastern Europe and provides valuable insights into the Middle Pleistocene terrestrial ecosystems of Europe, and hominin adaptations in particular. Full article
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